#  Evolution Faculty 

 



Through field, laboratory, and model-based studies along with the outstanding collections, facilities and other resources of Harvard’s natural history institutions—the Museum of Comparative Zoology, the Harvard University Herbaria, the Arnold Arboretum, and the Harvard Forest—we investigate the origins and maintenance of biological diversity.



 

  [### Colleen Cavanaugh

 ](/people/colleen-cavanaugh) <cavanaug@fas.harvard.edu>Edward C. Jeffrey Professor of Biology

Not Accepting Graduate Students for 2026-2027

 

 

 

      ![Colleen Cavanauth with medium dark hair wearing glasses, a black top and grey jacket in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/colleen_cavanaugh_crop.jpeg?itok=lYix3xo3) 

 

 

 

   [### Jeannine Cavender-Bares

 ](/people/jeannine-cavender-bares) <jcavender@fas.harvard.edu>Climate Action Acceleration Professor of Organismic and Evolutionary Biology

Director of the Harvard University Herbaria

Chair of the Harvard Forest Leadership Committee

 

 

 

      ![Jeannine Cavender-Bares smiling, with long dark hair, standing in a field](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/2025-04/Cavender-Bares_6_2025.jpg?itok=6tk4hyhg) 

 

 

 

   [### Charles C. Davis

 ](/people/charles-c-davis) <cdavis@oeb.harvard.edu>Professor of Organismic and Evolutionary Biology

Curator of Vascular Plants in the Harvard University Herbaria

 

 

 

      ![Charles Davis, with short brown hair wearing a suit outside in front of a tree](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/charles_davis_crop.jpeg?itok=iVPwX7wY) 

 

 

 

   [### Benjamin L. de Bivort

 ](/people/benjamin-de-bivort) <debivort@oeb.harvard.edu>Professor of Organismic and Evolutionary Biology

Co-Chair of the Department of Organismic and Evolutionary Biology

 

 

 

      ![Benjamin de Bivort with beard and mustache wearing a red shirt and black jacket smiling](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/benjamin_de_bivort-2.jpeg?itok=wrRgNY_x) 

 

 

 

   [### Michael M. Desai

 ](/people/michael-desai) <mdesai@oeb.harvard.edu>Fisher Professor of Natural History

Co-Director of the Systems Synthetic and Quantitative Biology PhD Program

 

 

 

      ![Michael Desai with short dark hair and beard, wearing a blue check button down, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/michael_desai_crop.jpeg?itok=I8_JgNYw) 

 

 

 

   [### Scott V. Edwards

 ](/people/scott-v-edwards) <sedwards@fas.harvard.edu>Professor of Organismic and Evolutionary Biology

Curator of Ornithology in the Museum of Comparative Zoology

 

 

 

      ![Scott Edwards smiling, with short grey hair and beard, wearing light blue button down, with light grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/scott_edwards_crop.jpeg?itok=CTYKumrd) 

 

 

 

   [### Cassandra G. Extavour

 ](/people/cassandra-g-extavour) <extavour@oeb.harvard.edu>Timken Professor of Organismic and Evolutionary Biology and of Molecular and Cellular Biology

Howard Hughes Medical Institute Investigator

 

 

 

      ![Cassandra Extavour with brown hair wearing a black blouse, in front of window](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/extavour.jpeg?itok=VQVxnSuq) 

 

 

 

   [### Brian D. Farrell

 ](/people/brian-d-farrell) <farrellb@oeb.harvard.edu>Monique and Philip Lehner Professor for the Study of Latin America

Professor of Biology

Curator of Entomology in the Museum of Comparative Zoology

 

 

 

      ![Brian D. Farrell with grey beard and hair, wearing white short sleeve button down, with spike green leaves behind him](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/farrell.jpg?itok=B9n0E1GD) 

 

 

 

   [### William (Ned) Friedman

 ](/people/william-friedman) <ned@oeb.harvard.edu>Arnold Professor of Organismic and Evolutionary Biology

Director of the Arnold Arboretum

Faculty Fellow of the Arnold Arboretum

 

 

 

      ![William Friedman smiling, with gray hair and glasses wearing a suit, in front of leafy trees](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/willian_friedman.jpg?itok=fGW9xW-w) 

 

 

 

   [### Gonzalo Giribet

 ](/people/gonzalo-giribet) <ggiribet@g.harvard.edu>Professor of Organismic and Evolutionary Biology

Curator of Invertebrate Zoology in the Museum of Comparative Zoology

Director of the Museum of Comparative Zoology

 

 

 

      ![Gonzalo Giribet sitting, with short hair, wearing white button down, in front of white wall with framed picture](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/gonzalo_giribet_2021.jpeg?itok=F_EdC_b4) 

 

 

 

   [### David A. Haig

 ](/people/david-haig) <dhaig@oeb.harvard.edu>George Putnam Professor of Organismic and Evolutionary Biology 

Associate of Dudley House

 

 

 

      ![David Haig with gray hair, a beard, wearing glasses and a suit, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/david_haig_crop.jpeg?itok=5h0-SS6i) 

 

 

 

   [### Daniel L. Hartl

 ](/people/daniel-l-hartl) <dhartl@oeb.harvard.edu>Higgins Professor of Biology

Not Accepting Graduate Students for 2026-2027

 

 

 

      ![Dan Hartl with gray hair and a beard wearing glasses and a blue shirt](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/dan_hartl_crop.jpeg?itok=rdmLVaR5) 

 

 

 

   [### Hopi E. Hoekstra

 ](/people/hopi-e-hoekstra) [hopi\_hoekstra@harvard.edu](mailto:hopi_hoekstra@harvard.edu)Edgerley Family Dean of the Faculty of Arts and Sciences

C.Y. Chan Professor of Arts and Sciences

Xiaomeng Tong and Yu Chen Professor of Life Sciences

Senior Fellow of the Society of Fellows, ex officio

Not Accepting Graduate Students

 

 

 

      ![Hopi Hoekstra smiling, learning on doorway, with long hair and black shirt with white pattern](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/hopi_hoekstra.jpeg?itok=iTl97Pr8) 

 

 

 

   [### Mandë Holford

 ](/people/mande-holford) <mholford@fas.harvard.edu>Professor of Organismic and Evolutionary Biology

Curator of Malacology in the Museum of Comparative Zoology

 

 

 

      ![Mande Holford, smiling, with long dark hair and blue and white patterned tops, in front of dark background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/mande_holford.jpg?itok=m6_FG0Zu) 

 

 

 

   [### Robin Hopkins

 ](/people/robin-hopkins) <rhopkins@fas.harvard.edu>Professor of Organismic and Evolutionary Biology

Faculty Fellow of the Arnold Arboretum

Head Tutor, Integrative Biology Undergraduate Program

 

 

 

      ![Robin Hopkins smiling, with short hair, grey sweater and pink top, in front of light grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/robin_hopkins_crop.jpeg?itok=WSizin16) 

 

 

 

   [### Elena Kramer

 ](/people/elena-kramer) <ekramer@oeb.harvard.edu>Bussey Professor of Organismic and Evolutionary Biology

Dean of Science Education

Faculty Fellow of the Arnold Arboretum

Co-Director of Harvard Integrated Life Sciences

Director of Graduate Programs in the Natural Sciences

 

 

 

      ![Elena Kramer smiling, with long dark hair, glasses, and wearing a blue shirt, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/elena_kramer_crop.jpeg?itok=gh8aGCM1) 

 

 

 

   [### George V. Lauder

 ](/people/george-v-lauder) <glauder@oeb.harvard.edu>Henry Bryant Bigelow Professor of Ichthyology

Curator of Ichthyology in the Museum of Comparative Zoology

 

 

 

      ![George Lauder with short gray hair wearing glasses and a plaid shirt in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/george_lauder_crop.jpeg?itok=SCZBahUU) 

 

 

 

   [### James Mallet

 ](/people/james-mallet) <jmallet@oeb.harvard.edu>Professor of Organismic and Evolutionary Biology in Residence

Associate of the Museum of Comparative Zoology

 

 

 

      ![James Mallet with short grey hair, wearing white shirt and dark jacket, with  grass and brick building behind him](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/jamesmallet_photobytonyrinaldo.jpeg?itok=S8FG4AhT) 

 

 

 

   [### Martin A. Nowak

 ](/people/martin-nowak) [martin\_nowak@harvard.edu](mailto:martin_nowak@harvard.edu)On Leave

Professor of Mathematics and Biology

 

 

 

      ![Martin A. Nowak wearing a suit and tie, in front of a chalkboard with equations on it](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/martin_nowak.jpg?itok=9ttO8Fd1) 

 

 

 

   [### Javier Ortega-Hernández

 ](/people/javier-ortega-hernandez) <jortegahernandez@fas.harvard.edu>Associate Professor of Organismic and Evolutionary Biology

Curator of Invertebrate Paleontology in the Museum of Comparative Zoology

Affiliate of the Department of Earth and Planetary Sciences

 

 

 

      ![Javier Ortega-Hernandez wearing colorful glasses, a pink check shirt, and with a short dark beard, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/javier_ortega-hernandez_crop.jpeg?itok=RFA7tedh) 

 

 

 

   [### Stephanie E. Pierce

 ](/people/stephanie-e-pierce) <spierce@oeb.harvard.edu>Professor of Organismic and Evolutionary Biology

Curator of Vertebrate Paleontology and Interim Curator of Mammalogy in the Museum of Comparative Zoology

Alexander Agassiz Professor of Zoology in the Museum of Comparative Zoology

Director of Graduate Studies

 

 

 

      ![Stephanie Pierce with long, blonde hair wearing a green sweater, patterned shirt, in front of grey background ](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/stephanie_pierce_crop.jpeg?itok=Nx5G7I5S) 

 

 

 

   [### Naomi E. Pierce

 ](/people/naomi-e-pierce) <npierce@oeb.harvard.edu>Sidney A. and John H. Hessel Professor of Biology

Curator of Lepidoptera in the Museum of Comparative Zoology

Senior Fellow of the Society of Fellows

 

 

 

      ![Naomi Pierce with long grey hair pulled back, glasses on head, dark sweater, light shirt, and colorful scarf](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/naomi_pierce_crop.jpeg?itok=ZQmyvtl1) 

 

 

 

   [### Pardis Sabeti

 ](/people/pardis-sabeti) <pardis@broadinstitute.org>Professor of Organismic and Evolutionary Biology

Professor of Immunology and Infectious Diseases

Institute Member, Broad Institute of MIT and Harvard

Howard Hughes Medical Institute Investigator

 

 

 

      ![Pardis Sabeti smiling, with long light brown hair and black shirt, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/sabeti.jpg?itok=T5Y7jqyj) 

 

 

 

   [### Mansi Srivastava

 ](/people/mansi-srivastava) <mansi@oeb.harvard.edu>Professor of Organismic and Evolutionary Biology

Curator of Invertebrate Zoology in the Museum of Comparative Zoology

Co-Chair of the Department of Organismic and Evolutionary Biology

 

 

 

      ![Mansi Srivistava with long, brown hair wearing a green blouse, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/mansi_srivastava_crop.jpeg?itok=GBxFocqa) 

 

 

 

   [### John Wakeley

 ](/people/john-wakeley) <wakeley@fas.harvard.edu>Professor of Organismic and Evolutionary Biology

 

 

 

      ![Professor John Wakeley with short hair and light shirt, standing in front of a tree](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/2025-08/Wakeley-John_Radcliffe%20copy.jpg?itok=vRbd_UnF)