#  Evolutionary Developmental Biology Faculty 

 



We seek to understand how changes in developmental programs have shaped the evolution of morphological diversity. Evolutionary developmental biology (EDB, but often called “Evo-Devo”) integrates traditional research on organismal evolutionary biology (systematics, paleontology, and comparative anatomy) with molecular embryology, genetics and genomics. EDB is a thriving new sub-discipline of biology with its own questions, approaches and methods. The field seeks to forge a unifying integration of genomic, developmental, organismal, population, paleontological and natural selection approaches to evolutionary change.



 

  [### Carrie Albertin

 ](/people/carrie-albertin) <calbertin@fas.harvard.edu>Assistant Professor of Organismic and Evolutionary Biology

Curator of Malacology in the Museum of Comparative Zoology

Accepting Graduate Students for the 2027-2028 Academic Year

 

 

 

      ![Carrie Albertin smiling, with long blonde hair and a blue and white top](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/2026-01/Carrie%20Albertin%20crop.jpeg?itok=2ki0DeFS) 

 

 

 

   [### Cassandra G. Extavour

 ](/people/cassandra-g-extavour) <extavour@oeb.harvard.edu>Timken Professor of Organismic and Evolutionary Biology and of Molecular and Cellular Biology

Howard Hughes Medical Institute Investigator

Accepting Graduate Students for the 2027-2028 Academic Year

 

 

 

      ![Cassandra Extavour with brown hair wearing a black blouse, in front of window](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/extavour.jpeg?itok=VQVxnSuq) 

 

 

 

   [### Gonzalo Giribet

 ](/people/gonzalo-giribet) <ggiribet@g.harvard.edu>Professor of Organismic and Evolutionary Biology

Curator of Invertebrate Zoology in the Museum of Comparative Zoology

Director of the Museum of Comparative Zoology

Accepting Graduate Students for the 2027-2028 Academic Year

 

 

 

      ![Gonzalo Giribet sitting, with short hair, wearing white button down, in front of white wall with framed picture](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/gonzalo_giribet_2021.jpeg?itok=F_EdC_b4) 

 

 

 

   [### Hopi E. Hoekstra

 ](/people/hopi-e-hoekstra) [hopi\_hoekstra@harvard.edu](mailto:hopi_hoekstra@harvard.edu)Edgerley Family Dean of the Faculty of Arts and Sciences

C.Y. Chan Professor of Arts and Sciences

Xiaomeng Tong and Yu Chen Professor of Life Sciences

Senior Fellow of the Society of Fellows, ex officio

Not Accepting Graduate Students

 

 

 

      ![Hopi Hoekstra smiling, learning on doorway, with long hair and black shirt with white pattern](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/hopi_hoekstra.jpeg?itok=iTl97Pr8) 

 

 

 

   [### Mandë Holford

 ](/people/mande-holford) <mholford@fas.harvard.edu>Professor of Organismic and Evolutionary Biology

Curator of Malacology in the Museum of Comparative Zoology

Accepting Graduate Students for the 2027-2028 Academic Year

 

 

 

      ![Mande Holford, smiling, with long dark hair and blue and white patterned tops, in front of dark background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/mande_holford.jpg?itok=m6_FG0Zu) 

 

 

 

   [### Elena Kramer

 ](/people/elena-kramer) <ekramer@oeb.harvard.edu>Bussey Professor of Organismic and Evolutionary Biology

Dean of Science Education

Faculty Fellow of the Arnold Arboretum

Co-Director of Harvard Integrated Life Sciences

Director of Graduate Programs in the Natural Sciences

Accepting Graduate Students for the 2027-2028 Academic Year

 

 

 

      ![Elena Kramer smiling, with long dark hair, glasses, and wearing a blue shirt, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/elena_kramer_crop.jpeg?itok=gh8aGCM1) 

 

 

 

   [### Javier Ortega-Hernández

 ](/people/javier-ortega-hernandez) <jortegahernandez@fas.harvard.edu>Associate Professor of Organismic and Evolutionary Biology

Curator of Invertebrate Paleontology in the Museum of Comparative Zoology

Affiliate of the Department of Earth and Planetary Sciences

Accepting Graduate Students for the 2027-2028 Academic Year

 

 

 

      ![Javier Ortega-Hernandez wearing colorful glasses, a pink check shirt, and with a short dark beard, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/javier_ortega-hernandez_crop.jpeg?itok=RFA7tedh) 

 

 

 

   [### Mansi Srivastava

 ](/people/mansi-srivastava) <mansi@oeb.harvard.edu>Professor of Organismic and Evolutionary Biology

Curator of Invertebrate Zoology in the Museum of Comparative Zoology

Co-Chair of the Department of Organismic and Evolutionary Biology

 

 

 

      ![Mansi Srivistava with long, brown hair wearing a green blouse, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/mansi_srivastava_crop.jpeg?itok=GBxFocqa)