# Population & Evolutionary Genetics Faculty We study variation within and between populations at the molecular and phenotypic levels. We strive to understand the evolutionary forces molding genetic variation in natural populations and the mechanistic basis of phenotypic evolution. Our study systems vary from microbes to animals and plants, and to mathematical theory. [### Scott V. Edwards ](/people/scott-v-edwards) Professor of Organismic and Evolutionary Biology Curator of Ornithology in the Museum of Comparative Zoology ![Scott Edwards smiling, with short grey hair and beard, wearing light blue button down, with light grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/scott_edwards_crop.jpeg?itok=CTYKumrd) [### Brian D. Farrell ](/people/brian-d-farrell) Monique and Philip Lehner Professor for the Study of Latin America Professor of Biology Curator of Entomology in the Museum of Comparative Zoology ![Brian D. Farrell with grey beard and hair, wearing white short sleeve button down, with spike green leaves behind him](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/farrell.jpg?itok=B9n0E1GD) [### Daniel L. Hartl ](/people/daniel-l-hartl) Higgins Professor of Biology Not Accepting Graduate Students for 2026-2027 ![Dan Hartl with gray hair and a beard wearing glasses and a blue shirt](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/dan_hartl_crop.jpeg?itok=rdmLVaR5) [### Hopi E. Hoekstra ](/people/hopi-e-hoekstra) [hopi\_hoekstra@harvard.edu](mailto:hopi_hoekstra@harvard.edu)Edgerley Family Dean of the Faculty of Arts and Sciences C.Y. Chan Professor of Arts and Sciences Xiaomeng Tong and Yu Chen Professor of Life Sciences Senior Fellow of the Society of Fellows, ex officio Not Accepting Graduate Students ![Hopi Hoekstra smiling, learning on doorway, with long hair and black shirt with white pattern](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/hopi_hoekstra.jpeg?itok=iTl97Pr8) [### Robin Hopkins ](/people/robin-hopkins) Professor of Organismic and Evolutionary Biology Faculty Fellow of the Arnold Arboretum Head Tutor, Integrative Biology Undergraduate Program ![Robin Hopkins smiling, with short hair, grey sweater and pink top, in front of light grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/robin_hopkins_crop.jpeg?itok=WSizin16) [### Elena Kramer ](/people/elena-kramer) Bussey Professor of Organismic and Evolutionary Biology Dean of Science Education Faculty Fellow of the Arnold Arboretum Co-Director of Harvard Integrated Life Sciences Director of Graduate Programs in the Natural Sciences ![Elena Kramer smiling, with long dark hair, glasses, and wearing a blue shirt, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/elena_kramer_crop.jpeg?itok=gh8aGCM1) [### James Mallet ](/people/james-mallet) Professor of Organismic and Evolutionary Biology in Residence Associate of the Museum of Comparative Zoology ![James Mallet with short grey hair, wearing white shirt and dark jacket, with grass and brick building behind him](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/jamesmallet_photobytonyrinaldo.jpeg?itok=S8FG4AhT) [### Martin A. Nowak ](/people/martin-nowak) [martin\_nowak@harvard.edu](mailto:martin_nowak@harvard.edu)On Leave Professor of Mathematics and Biology ![Martin A. Nowak wearing a suit and tie, in front of a chalkboard with equations on it](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/martin_nowak.jpg?itok=9ttO8Fd1) [### Pardis Sabeti ](/people/pardis-sabeti) Professor of Organismic and Evolutionary Biology Professor of Immunology and Infectious Diseases Institute Member, Broad Institute of MIT and Harvard Howard Hughes Medical Institute Investigator ![Pardis Sabeti smiling, with long light brown hair and black shirt, in front of grey background](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/oeb/files/sabeti.jpg?itok=T5Y7jqyj) [### John Wakeley ](/people/john-wakeley) Professor of Organismic and Evolutionary Biology ![Professor John Wakeley with short hair and light shirt, standing in front of a tree](/sites/g/files/omnuum6811/files/styles/hwp_4_5__690x865/public/2025-08/Wakeley-John_Radcliffe%20copy.jpg?itok=vRbd_UnF)